I think Link is trying to say is that the more timid Salmon live longer, and have a better life. So Darwin's theory on Natural section is opposite when it comes to Salmon in this story. That's why Link also says survival of the timid, because the timid fish will not try to go up stream like the other fishes that think they can do it. The fish that try die fast, and the fish that just stay at the bottom of dam actually get to spawn. So in the end the timid are the best survivals not the fittest.
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